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密度对四季竹地上生物量分配格局及异速增长模式的制约性调节

郭子武,杨清平,李迎春,陈双林**   

  1. (中国林业科学研究院亚热带林业研究所, 浙江富阳 311400)
  • 出版日期:2013-03-10 发布日期:2013-03-10

Restrictive regulation of stand density on aboveground biomass allocation and allometric pattern of Oligostachyum lubricum. 

GUO Zi-wu, YANG Qing-ping, LI Ying-chun, CHEN Shuang-lin**   

  1. (Research Institute of Subtropical Forestry, Chinese Academy of Forestry, Fuyang 311400, Zhejiang, China)
  • Online:2013-03-10 Published:2013-03-10

摘要: 为了给四季竹(Oligostachyum lubricum)高效培育的林分结构建立提供理论依据,对立竹胸径基本一致的4种立竹密度(24600~29800株·hm-2,D1;37500~42600株·hm-2,D2;46500~52800 株·hm-2,D3;76500~85500株·hm-2,D4)四季竹纯林进行了1~3 a立竹地上生物量积累、分配与异速生长模式的研究。结果表明:四季竹立竹构件生物量分配比例秆>叶>枝。随着立竹年龄增大,不同密度的四季竹林立竹秆、枝、叶生物量和地上生物量及叶/枝、叶/秆、枝/秆构件生物量比总体上均呈增大趋势,且2 a、3 a立竹显著高于1 a立竹,枝、叶生物量分配比例呈升高趋势,而秆生物量分配比例呈下降趋势。随着立竹密度增大,1~3 a立竹地上生物量、构件生物量总体上呈“∧”型变化,D1~D3密度时逐渐升高,D4密度时显著下降,各年龄立竹枝、叶生物量分配比例降低,而秆生物量比例增大,叶/枝、叶/秆、枝/秆生物量比总体上1 a立竹呈倒“N”型变化,2 a、3 a立竹呈下降趋势。叶-枝构件生物量符合近等速增长模式,异速生长指数随密度的增大而小幅度下降,叶-秆、枝-秆构件生物量符合简单异速增长模式,异速生长指数随密度的增大分别呈升高、先升高后降低的变化趋势。研究表明,当超出一定密度时(D2密度以上),四季竹立竹生物量分配更趋向于支撑构件(秆),以促进立竹纵向生长来获取更多的光资源;46500~52800株·hm-2是试验四季竹林立竹生物量高效积累和有效分配的密度。

关键词: FACE, C4植物, 稗草, 光合作用, 适应, 气孔导度

Abstract: In order to provide a theoretical basis for establishing a high-productive Oligostachyum lubricum stand, this paper studied the aboveground biomass accumulation, allocation and the allometric pattern of 1-3 yearold O. lubricum at four stand densities (24600-29800 stems·hm-2, D1; 37500-42600 stems·hm-2, D2; 46500-52800 stems·hm-2, D3; and 76500-85500 stems·hm-2, D4) with basically the same DBH. The allocation proportion of the O. lubricum modular biomass was in the order stem>branch>leaf. With the increasing age of the O. lubricum, the modular biomass and aboveground biomass as well as the leaf/branch, leaf/stem, and branch/stem ratios at the four stand densities all had an increasing trend, and were significantly higher for 2 and 3-year-old O. lubricum than for 1-year-old O. lubricum. The allocation proportion of the branch and leaf biomass presented an increasing trend, while that of the stem biomass was in adverse. With the increasing stand density, the modular biomass and aboveground biomass of 1-3-year-old O. lubricum were overall in “∧” shape, i.e., increased gradually at stand densities D1-D3 but decreased significantly at stand density D4. The allocation proportion of the leaf and branch biomass of 1-3-year-old O. lubricum decreased, while that of the stem biomass increased. The leaf/branch, leaf/stem, and branch/stem ratios of 1-year-old O. lubricum were in inverse “N” shape, while those of 2- and 3-year-old decreased on the whole. The leafbranch biomass accorded with near-constant growth model, and the allometric index decreased slightly with increasing stand density; whereas the leaf-stem biomass and the branch-stem biomass accorded with simple allometric model, and their allometric indices presented a trend of increasing and of decreased after an initial increase with the increase of the stand density, respectively. All the results indicated that when the stand density was higher than 37500-42600 stems·hm-2, more biomass was tended to allocate in stem to promote the longitudinal growth to obtain more light resource, and the suitable stand density for the high-efficient biomass accumulation and allocation was 46500-52800 stems·hm-2.