1Key Laboratory of Genetic Breeding and Aquaculture Biology of Freshwater Fishes, Ministry of Agriculture, Freshwater Fisheries Research Center, Chinese Academy of Fishery Sciences, Wuxi 214081, Jiangsu, China; 2Extension Station of Aquiculture Technology of Jiangyin, Jiangyin 214431, JIangsu, China;3Research Center for the Yangtze River Rare Fish Conservation, Jiangyin 214431, Jiangsu, China