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空间格局和物种组合对三个植物种行为的影响

张作亮;马银山;赵川;李传龙;张世挺   

  1. 兰州大学干旱与草地生态教育部重点实验室, 兰州 730000
  • 收稿日期:2008-06-18 修回日期:1900-01-01 出版日期:2009-03-10 发布日期:2009-03-10

Effects of spatial pattern and species combination on the performance of three plant species.

ZHANG Zuo-liang;MA Yin-shan;ZHAO Chuan;LI Chuan-long;ZHANG Shi-ting   

  1. Key Laboratory of Arid and Grassland Agroecology of Ministry of Education, Lanzhou University, Lanzhou 730000, China
  • Received:2008-06-18 Revised:1900-01-01 Online:2009-03-10 Published:2009-03-10

摘要: 以青藏高原东部高寒草甸弃耕地种子大小不同的3个常见物种:2个1年生物种鼬瓣花(Galeopsis bifida)和菊叶香藜(Chenopodium foetidum)及1个2年生物种四数獐牙菜(Swertia tetraptera)为试验材料,通过单播、2个、3个物种的混播,分别以2个密度(高密度和低密度),按照随机或者种内积聚的空间格局形式播种,研究空间格局、物种组合对不同种子大小植物行为的影响。结果表明:1)空间格局和物种组合对3物种的出苗率没有显著影响,但在一定程度上影响着物种竞争能力等级,大种子鼬瓣花在种间竞争中占据着优势地位,小种子菊叶香藜在种内集聚格局下获益;四数獐牙菜的种子大小介于二者之间,其种间竞争的结果表现复杂;2)集聚格局下菊叶香藜的幼苗死亡率低,地上生物量比随机格局下显著高;集聚格局下鼬瓣花死亡率高,通过密度-格局作用,高密集聚下的地上生物量才表现显著较低;3)与其他物种混播相比,鼬瓣花单播下死亡率高,地上生物量低;当混播中存在四数獐牙菜时,菊叶香藜的死亡率比单播下高,物种组合对其地上生物量没有显著影响;四数獐牙菜与鼬瓣花混播时死亡率最低,3个物种混播时其死亡率最高。

关键词: 硫酸盐纸浆废液, 木素, 酚醛树脂, PAPI, 红外光谱分析

Abstract: Taking the common three plant species with different seed size (two an nuals Galeopsis bifida and Chenopodium foetidum and a biennial Swertia tetraptera) in abandoned fields of eastern Tibetan Plateau as test materials, this paper studied their performance under effects of different spatial pattern (aggregated vs. random distribution, and low vs. high density) and species combination (mon oculture vs. mix-culture). The results showed that spatial pattern and species c ombination had no significant effects on the emergence rate, but affected the co mpetition hierarchy of the three species to some degree. Large-seed species G . bifida exhibited dominance in interspecific competition, small-seed species C. foetidum benefited from intraspecific aggregated pattern, while S. tet raptera, whose seed size was between that of G. bifida and C. foetidum , showed complexity in interspecific competition. For C. foetidum, its seedl ing mortality rate in aggregated pattern was lower, and its aboveground biomass was significantly higher in aggregated pattern than in random pattern. For G. bifida, its seedling mortality rate in aggregated pattern was higher, and its aboveground biomass was signif icantly lower in high density and aggregated pattern. The seedling mortality rat e of G. bifida was higher and its aboveground biomass was lower in monocultu re t han in mix-culture. For C. foetidum, its seedling mortality rate was higher in m ix-culture with S. tetraptera than in monoculture, but there was no signifi cant difference in its aboveground biomass among different species combination. For S. tetraptera, its seedling mortality rate was the lowest in mix-culture with G. bifida, but the highest in mix-culture with G. bifida and C. foetidum.

Key words: Sulfate paper pulp waste liquid, Lignin, Phenol formaldehyde resin, Polyfunctional aqueous polymer isocyanate, Infrared spectrum analysis